akfg1.jpg (41982 bytes)   Emerging Area of Aging Research:
  Long-lived Animals with
  "Negligible Senescence"
Research 1-8
Research 9-14
Publications on Long Lived Animals
Principal Investigators



Leonard Hayflick, Ph.D.
Professor of Anatomy University of California, San Francisco

Relationship to the Project:
Dr. Hayflick, best known for the Hayflick Limit showing that cultured normal cells have a limited capacity for replication, wrote the popular book, “How and Why We Age”, in August 1994. Mr. Guerin read the book with interest a year later. He especially noted the chapter entitled “Some Animals Age, Some Do Not”. It was the first time Mr. Guerin had read this information from a scientific source, although later he came across detailed information on this phenomenon coined “negligible senescence” by Caleb Finch at USC. Mr. Guerin started the project that came to be known as the Centenarian Species and Rockfish Project in August 1995 as a result of this information.

Later on in 1995 the Director met Dr. Hayflick at a Gerontological Society of America meeting. Mr. Guerin told him of his background in Project Management, and his interest in assisting research in this area. He asked Dr. Hayflick who was researching these long-lived animals, and the reply was “Nobody is, but they should be!” From this point forward Dr. Hayflick has served as a Scientific Advisor to this project. His biography is below

His biography is available on Wikipedia

Aubrey D.N.J. de Grey


Name: Dr. Aubrey D.N.J. de Grey


Current post: Chief Science Officer, SENS Research Foundation

Research area: the role and causes of all the accumulating and eventually pathogenic molecular and cellular side-effects of metabolism that constitute mammalian aging; the design of interventions to reverse and/or obviate this accumulation.

Bio paragraph (e.g., if you have invited me to speak and are creating promotional material):

Dr. Aubrey de Grey is a biomedical gerontologist based in Cambridge, UK and Mountain View, California, USA, and is the Chief Science Officer of SENS Research Foundation, a California-based 501(c)(3) charity dedicated to combating the aging process. He is also Editor-in-Chief of Rejuvenation Research, the world’s highest-impact peer-reviewed journal focused on intervention in aging. He received his BA and Ph.D. from the University of Cambridge in 1985 and 2000 respectively. His original field was computer science, and he did research in the private sector for six years in the area of software verification before switching to biogerontology in the mid-1990s. His research interests encompass the characterisation of all the accumulating and eventually pathogenic molecular and cellular side-effects of metabolism (“damage”) that constitute mammalian aging and the design of interventions to repair and/or obviate that damage. He has developed a possibly comprehensive plan for such repair, termed Strategies for Engineered Negligible Senescence (SENS), which breaks aging down into seven major classes of damage and identifies detailed approaches to addressing each one. A key aspect of SENS is that it can potentially extend healthy lifespan without limit, even though these repair processes will probably never be perfect, as the repair only needs to approach perfection rapidly enough to keep the overall level of damage below pathogenic levels. Dr. de Grey has termed this required rate of improvement of repair therapies “longevity escape velocity”. Dr. de Grey is a Fellow of both the Gerontological Society of America and the American Aging Association, and sits on the editorial and scientific advisory boards of numerous journals and organisations.

Tel.: +44 1223 366197 or +1 650 938 6100

Email: aubrey@sens.org

Date of birth: 20th April, 1963

Nationality: British

Marital status: Married to Dr. Adelaide T.C. Carpenter; no children


  B.A. (1985), M.A. (1988) and Ph.D. (2000), University of Cambridge, Cambridge, UK

Past employment:

  1985-1986: Sinclair Research, Cambridge, UK

  1986-1992: Man-Made Minions, Cambridge, UK

  1992 (Jan-Oct): EO Computer, Cambridge, UK

  1992-2006: Department of Genetics, University of Cambridge, Cambridge, UK

  2003-2009: Chairman and Chief Science Officer, Methuselah Foundation


  2003: Gerontological Society of America: Fellow

  2004: World Transhumanist Association: H.G. Wells Award

  2005: American Aging Association: Fellow


Society memberships:

  American Aging Association (member of board of directors, 2002-2007 and 2010-present)

  International Association of Biomedical Gerontology (chair, 2003-2005)

  British Society for Research on Ageing (member of board of directors, 2003-2006)

  Gerontological Society of America

  International Coenzyme Q10 Association

  Mitochondrion Research Society


Scientific advisory boards (unpaid):

  Maximum Life Foundation

  Legendary Pharmaceuticals

  Centenarian Species and Rockfish Project

  Alcor Life Extension Foundation

  Singularity Institute for Artificial Intelligence

  Foresight Institute


  Lifeboat Foundation


  Campaign for Aging Research


Conferences organised:

  Euromit 4, Cambridge, UK, September 1999 (co-organiser)

  Strategies for Engineered Negligible Senescence, 1st roundtable, Oakland, CA, USA, October 2000

  Strategies for Engineered Negligible Senescence, 2nd roundtable, Los Angeles, CA, USA, Aug. 2001

  Strategies for Engineered Negligible Senescence, 3rd roundtable, Cambridge, UK, December 2002

  International Association of Biomedical Gerontology, 10th Congress, Cambridge, UK, Sept. 2003

  Strategies for Engineered Negligible Senescence, 4th roundtable, Bethesda, MD, USA, July 2004

  Strategies for Engineered Negligible Senescence, 2nd conference, Cambridge, UK, September 2005

  Strategies for Engineered Negligible Senescence, 3rd conference, Cambridge, UK, September 2007

  Understanding Aging: Biomedical and Bioenginering Approaches, Los Angeles, USA, June 2008

  Strategies for Engineered Negligible Senescence, 4th conference, Cambridge, UK, September 2009

  Strategies for Engineered Negligible Senescence, 5th conference, Cambridge, UK, September 2011

  Strategies for Engineered Negligible Senescence, 6th conference, Cambridge, UK, September 2013


Journal editorial board memberships:

  Rejuvenation Research (editor-in-chief)

  Disruptive Science and Technology (executive editor)

  Mitochondrion (associate editor)

  AGE – Journal of the American Aging Association

  Current Stem Cell Research & Therapy

  Current Gerontology and Geriatrics Research



- Articles in peer-reviewed journals:

de Grey ADNJ. A proposed refinement of the mitochondrial free radical theory of aging. BioEssays 1997; 19(2):161-166.

de Grey ADNJ. A mechanism proposed to explain the rise in oxidative stress during aging. J Anti-Aging Med 1998; 1(1):53-66.

de Grey ADNJ. Incorporation of transmembrane hydroxide transport into the chemiosmotic theory. Bioelectrochem Bioenerg 1999; 49(1-2):43-50.

de Grey ADNJ. The non-correlation between maximum longevity and enzymatic antioxidant levels among homeotherms; implications for retarding human aging. J Anti-Aging Med 2000; 3(1):25-36.

de Grey ADNJ. The reductive hotspot hypothesis: an update. Arch Biochem Biophys 2000; 373(1):295-301.

de Grey ADNJ. Mitochondrial gene therapy: an arena for the biomedical use of inteins. Trends Biotechnol 2000; 18(9):394-399.

de Grey ADNJ. A proposed mechanism for the lowering of mitochondrial electron leak by caloric restriction. Mitochondrion 2001; 1(2):129-139.

de Grey ADNJ. UK research on the biology of aging. Exp Gerontol 2001; 37(1):1-7.

de Grey ADNJ, Ames BN, Andersen JK, Bartke A, Campisi J, Heward CB, McCarter RJM, Stock G. Time to talk SENS: critiquing the immutability of human aging. Annals NY Acad Sci 2002; 959:452-462.

de Grey ADNJ. The reductive hotspot hypothesis of mammalian aging: membrane metabolism magnifies mutant mitochondrial mischief. Eur J Biochem 2002; 269(8):2003-2009.

de Grey ADNJ. HO2: the forgotten radical. DNA Cell Biol 2002; 21(4):251-257.

de Grey ADNJ, Baynes JW, Berd D, Heward CB, Pawelec G, Stock G. Is human aging still mysterious enough to be left only to scientists? BioEssays 2002; 24(7):667-676.

de Grey ADNJ. Bioremediation meets biomedicine: therapeutic translation of microbial catabolism to the lysosome. Trends Biotechnol 2002; 20(11):452-455.

de Grey ADNJ. An engineer’s approach to the development of real anti-aging medicine. Science’s SAGE KE 2003; 2003(1):vp1.

de Grey ADNJ. Challenging but essential targets for genuine anti-ageing drugs. Expert Opin Therap Targets 2003; 7(1):1-5.

de Grey ADNJ. Critique of the demographic evidence for “late-life non-senescence”. Biochem Soc Trans 2003; 31(2):452-454.

de Grey ADNJ. UK research on the biology of aging – the next ten years. Lifespan 2003; 11(1):1-4.

de Grey ADNJ. A hypothesis for the minimal overall structure of the mammalian plasma membrane redox system. Protoplasma 2003; 221(1-2):3-9.

de Grey ADNJ. The foreseeability of real anti-aging medicine: focusing the debate. Exp Gerontol 2003; 38(9):927-934.

de Grey ADNJ. Falsifying falsifications: the most critical task of theoreticians in biology. Med Hypoth 2004; 62(6):1012-1020.

de Grey ADNJ. Escape velocity: why the prospect of extreme human life extension matters now.  PLoS Biol 2004; 2(6):723-726.

de Grey ADNJ, Campbell FC, Dokal I, Fairbairn LJ, Graham GJ, Jahoda CAB, Porter ACG. Total deletion of in vivo telomere elongation capacity: an ambitious but possibly ultimate cure for all age-related human cancers. Annals NY Acad Sci 2004; 1019:147-170.

de Grey ADNJ. Biogerontologists’ duty to discuss timescales publicly. Annals NY Acad Sci 2004; 1019:542-545.

de Grey ADNJ. Inter-species therapeutic cloning: the looming problem of mitochondrial DNA and two possible solutions. Rejuvenation Res 2004; 7(2):95-98.

de Grey ADNJ. The unfortunate influence of the weather on the rate of aging: why human caloric restriction or its emulation may only extend life expectancy by 2-3 years. Gerontology 2005; 51(2):73-82.

de Grey ADNJ. Forces maintaining organellar genomes: is any as strong as genetic code disparity or hydrophobicity? BioEssays 2005; 27(4):436-446.

de Grey ADNJ. Whole-body interdiction of lengthening of telomeres: a proposal for cancer prevention. Front Biosci 2005; 10:2420-2429.

de Grey ADNJ. Reactive oxygen species production in the mitochondrial matrix: implications for the mechanism of mitochondrial mutation accumulation. Rejuvenation Res 2005; 8(1):13-17.

de Grey ADNJ. Resistance to debate on how to postpone ageing is delaying progress and costing lives. EMBO Rep 2005; 6(S1):S49-S53.

de Grey ADNJ, Alvarez PJJ, Brady RO, Cuervo AM, Jerome WG, McCarty PL, Nixon RA, Rittmann BE, Sparrow JR. Medical bioremediation: prospects for the application of microbial catabolic diversity to aging and several major age-related diseases. Ageing Res Rev 2005; 4(3):315-338.

de Grey ADNJ. Life extension, human rights, and the rational refinement of repugnance. J Med Ethics 2005; 31(11):659-663.

de Grey ADNJ. A strategy for postponing aging indefinitely. Stud Health Technol Inform 2005; 118:209-219.

de Grey ADNJ. The plasma membrane redox system: a candidate source of aging-related oxidative stress. AGE J Am Aging Assoc 2005; 27(2):129-138.

de Grey ADNJ. Appropriating microbial catabolism: a proposal to treat and prevent neurodegeneration. Neurobiol Aging 2006; 27(4):589-595.

de Grey ADNJ. Extrapolaholics Anonymous: why demographers’ rejections of a huge rise in cohort life expectancy in this century are overconfident. Annals NY Acad Sci 2006; 1067:83-93.

de Grey ADNJ. Foreseeable pharmaceutical repair of age-related extracellular damage. Curr Drug Targets 2006; 7(11):1469-1477.

de Grey ADNJ. Free radicals in aging: causal complexity and its biomedical implications. Free Radic Res 2006; 40(12):1244-1249.

de Grey ADNJ. The urgency dilemma: is life extension research a temptation or a test? Update 2006; 21(1):6-10.

de Grey ADNJ. The natural biogerontology portfolio: “defeating aging” as a multi-stage ultra-Grand Challenge. Annals NY Acad Sci 2007; 1100:409-423.

de Grey ADNJ. Protagonistic pleiotropy: why cancer may be the only pathogenic effect of accumulating nuclear mutations and epimutations in aging. Mech Ageing Dev 2007; 128(7-8):456-459.

Phoenix CR, de Grey ADNJ. A model of aging as accumulated damage matches observed mortality patterns and predicts the life-extending effects of prospective interventions. AGE 2007; 29(4):133-189.

de Grey ADNJ. Calorie restriction, post-reproductive lifespan and programmed aging: a plea for rigour. Annals NY Acad Sci 2007; 1119:296-305.

de Grey ADNJ. Alzheimer’s, atherosclerosis, and aggregates: A role for bacterial degradation. Nutr Rev 2007; 65(12 Pt. 2):S221-S227.

de Grey ADNJ. Life span extension research and public debate: societal considerations. Studies in Ethics, Law and Technology 2007; 1(1):5.

de Grey ADNJ. Our right to life. J Evol Technol 2008; 17(2):53-57.

de Grey ADNJ. The singularity and the Methuselarity: similarities and differences. Stud Health Technol Inform 2009; 149:195-202.

de Grey ADNJ. Making sure that health and wealth keep pace with extended life expectancy. United Academics Magazine 2012; 2012(April/May): http://www.united-academics.org/magazine/17748/making-sure-that-health-and-wealth-keep-pace-with-extended-life-expectancy/

de Grey ADNJ. Zeno’s paradox and the faith that technological game-changers are impossible. Gerontology 2012, in press.

Zealley BJ, de Grey ADNJ. Strategies for engineered negligible senescence (SENS). Gerontology 2012, in press.


- Reviews and commentaries:

de Grey ADNJ. More on mitochondria and senescence: Response to Gershon. BioEssays 1997; 19(6):534-534.

de Grey ADNJ. Review of Cadenas E and Packer L (eds), “Understanding the Process of Aging: The Roles of Mitochondria, Free Radicals, and Antioxidants”. J Anti-Aging Med 1999; 2(3):291.

de Grey ADNJ. Review of Harman D, Holliday R and Meydani M (eds), “Towards Prolongation of the Healthy Life Span: Practical Approaches to Intervention”. J Anti-Aging Med 1999; 2(3):292.

de Grey ADNJ. Popular gerontology comes of age. Review of Kirkwood TBL, “Time of our Lives”. BioEssays 1999; 21(9):802-803.

de Grey ADNJ. Biologists abandon Popper at their peril. BioEssays 2000; 22(2):206-207.

de Grey ADNJ. The function of mitochondrial dysfunction in aging. J Anti-Aging Med 2000; 3(1):109-112.

de Grey ADNJ. Review of Asard H, Berczi A and Caubergs RJ (eds), “Plasma Membrane Redox Systems and Their Role in Biological Stress and Disease”. Antioxid Redox Signal 2000; 2(2):371-372.

de Grey ADNJ. Redox 2000: the 5th international conference on plasma membrane redox systems and their role in biological stress and disease. Antioxid Redox Signal 2000; 2(2):373-374.

de Grey ADNJ. Review of Scheffler I, “Mitochondria”. J Am Aging Assoc 2000; 23(4):255.

de Grey ADNJ. Antioxidants and redox signaling: Internet resources. Antioxid Redox Signal 2000; 2(4):937-940.

de Grey ADNJ. Gerontologists and the media: the dangers of over-pessimism. Biogerontology 2000; 1(4):369-370.

de Grey ADNJ. Review of Wickens A, “The Causes of Aging”. Exp Gerontol 2000; 34(8):1017-1018.

de Grey ADNJ. Response to “telomere shortening with aging in human liver”. J Gerontol A Biol Sci Med Sci 2001; 56(6):B237-B238.

de Grey ADNJ. Response to “approaches and limitations to gene therapy for mitochondrial diseases,” Antioxid. Redox Signal. 2001;3:451-460. Antioxid Redox Signal 2001; 3(6):1153-1155.

de Grey ADNJ. Review of Finch CE and Kirkwood TBL, “Chance, development, and aging”. Exp Gerontol 2001; 36(10):1783-1785.

de Grey ADNJ. Nature of the aging process - Open discussion. Annals NY Acad Sci 2002; 959:463-465.

de Grey ADNJ, Gavrilov L, Olshansky SJ, Coles LS, Cutler RG, Fossel M, Harman SM. Antiaging technology and pseudoscience. Science 2002; 296(5568):656.

de Grey ADNJ. Three detailed hypotheses implicating oxidative damage to mitochondria as a major driving force in homeotherm aging. Eur J Biochem 2002; 269(8):1995.

de Grey ADNJ. Review of Nicholls DG and Ferguson SJ, “Bioenergetics 3”. Mitochondrion 2002; 2(3):211-213.

de Grey ADNJ. Fear of misrepresentation cannot justify silence about foreseeable life-extension biotechnology. BioEssays 2003; 25(1):94-95.

de Grey ADNJ. Overzealous maximum-likelihood fitting falsely convicts the slope heterogeneity hypothesis. Exp Gerontol 2003; 38(8):921-923.

de Grey ADNJ. Review of Mattson MP and Van Zant G (eds), “Stem Cells: A Cellular Fountain of Youth”. Exp Gerontol 2003; 38(9):1025-1026.

de Grey ADNJ. Breathtaking biomedical blind spots. Review of Guarente L, “Ageless Quest”. BioEssays 2004; 26(1):108-109.

de Grey ADNJ. Models on trial: falsifying overstated claims of generality does not falsify correctly-stated ones. Exp Gerontol 2004; 39(3):453.

de Grey ADNJ. Mitochondria in homeotherm aging: will detailed mechanisms consistent with the evidence now receive attention? Aging Cell 2004; 3(2):77.

de Grey ADNJ.  Report of open discussion on the future of life extension research. Annals NY Acad Sci 2004; 1019:552-553.

de Grey ADNJ. Welcome to Rejuvenation Research. Rejuvenation Res 2004; 7(1):1-2.

de Grey ADNJ. Leon Kass: quite substantially right. Rejuvenation Res 2004; 7(2):89-91.

de Grey ADNJ. Three self-evident life-extension truths. Rejuvenation Res 2004; 7(3):165-167.

de Grey ADNJ. Mitochondrial mutations in mammalian aging: an over-hasty about-turn? Rejuvenation Res 2004; 7(3):171-174.

de Grey ADNJ. Meeting report: Regenerate 2004: tissue engineering the human body, June 10-12, 2004, Seattle. Rejuvenation Res 2004; 7(3):226-228.

de Grey ADNJ. Aging, childlessness, or overpopulation: the future’s right to choose. Rejuvenation Res 2004; 7(4):237-238.

de Grey ADNJ. Meeting report: 8th International Symposium on the Maillard Reaction. Charleston, South Carolina, August 28-September 1 2004. Rejuvenation Res 2004; 7(4):257-260.

de Grey ADNJ. When and where to publish important findings: a casualty of biogerontology’s rise to respectability. Rejuvenation Res 2005; 8(1):1-2.

de Grey ADNJ. “The rate of aging”: a counterproductively undefinable term. Rejuvenation Res 2005; 8(2):77-78.

de Grey ADNJ. The ethical status of efforts to postpone aging: a reply to Hurlbut. Rejuvenation Res 2005; 8(3):129-130.

de Grey ADNJ. The SENS Challenge: $20,000 says the foreseeable defeat of aging is not laughable. Rejuvenation Res 2005; 8(4):207-210.

de Grey ADNJ. Meeting report: International Association of Biomedical Gerontology, 11th Congress, 13th-16th August, 2005, Aarhus, Denmark. Rejuvenation Res 2005; 8(4):264-266.

de Grey ADNJ. Like it or not, life extension research extends beyond biogerontology. EMBO Reports 2005; 6(11):1000.

de Grey ADNJ. Compression of morbidity: the hype and the reality, part 1. Rejuvenation Res 2006; 9(1):1-2.

de Grey ADNJ. Compression of morbidity: the hype and the reality, part 2. Rejuvenation Res 2006; 9(2):167-168.

de Grey ADNJ. Has Hippocrates had his day? Rejuvenation Res 2006; 9(3):371-373.

de Grey ADNJ. SENS survives the Challenge; now let’s get to work. Rejuvenation Res 2006; 9(4):429-430.

de Grey ADNJ. Is SENS a farrago? Rejuvenation Res 2006; 9(4):436-439.

de Grey ADNJ. SENS is hard, yes, but not too hard to try: a reply to Warner. Rejuvenation Res 2006; 9(4):443-445.

de Grey ADNJ. Aging and airborne HIV: a reassuring analogy. Rejuvenation Res 2007; 10(1):1-3.

de Grey ADNJ. New developments at Rejuvenation Research: high impact factor, advance online publication and more. Rejuvenation Res 2007; 10(2):125-126.

de Grey ADNJ. The case for prioritising research on late-onset life-extension interventions in mammals. Rejuvenation Res 2007; 10(3):257-259.

de Grey ADNJ. Edmonton: a future center for pioneering biomedical gerontology? Rejuvenation Res 2007; 10(3):345-347.

de Grey ADNJ. The need to debalkanize gerontology: a case study. Rejuvenation Res 2007; 10(4):431-434.

de Grey ADNJ. Understanding and tackling aging: two fields communicating (a little) at last. Rejuvenation Res 2007; 10(4):637-640.

de Grey ADNJ. Old people are people too: why it is our duty to fight aging to the death. Cato Unbound 2007(12): lead essay.

de Grey ADNJ. Curiosity is addictive, and this is not necessarily a good thing. Rejuvenation Res 2008; 11(1):1-3.

de Grey ADNJ. Man, machines, manufacturing and maintenance: merits of a much-maligned metaphor. Rejuvenation Res 2008; 11(2):277-279.

de Grey ADNJ. Long live the unreasonable man. Rejuvenation Res 2008; 11(3):541-542.

de Grey ADNJ. Combating the Tithonus Error: what works? Rejuvenation Res 2008; 11(4):713-715.

de Grey ADNJ. Trans-simianism and truthiness: hints of progress in the debate on whether aging is good. Rejuvenation Res 2008; 11(5):857-859.

de Grey ADNJ. Estimating rates of future biomedical progress: a reply to Bains. Biosci Hyp 2008; 1(5):286.

de Grey ADNJ. Aging: a foreseeable target of stem cells and regenerative medicine. World Stem Cell Report 2008:17-19.

de Grey ADNJ. Cautionary tales of the unexpected. Rejuvenation Res 2008; 11(6):989-990.

de Grey ADNJ. Consolidating the mission to defeat aging: a big step forward. Rejuvenation Res 2008; 11(6):1061-1063.

de Grey ADNJ. Cracks in social gerontology’s pro-aging edifice. Rejuvenation Res 2009; 12(1):1-3.

de Grey ADNJ. The curiously misunderstood role of evidence in designing new technology. Rejuvenation Res 2009; 12(2):75-76.

de Grey ADNJ. Why are so many people so unashamedly selfish about the prospect of combating aging? Rejuvenation Res 2009; 12(3):165-167.

de Grey ADNJ. How is mutant mitochondrial DNA clonally amplified? Much new evidence, still no answers. Rejuvenation Res 2009; 12(3):217-219.

de Grey ADNJ. Persons of tenure: less fortunate than they may seem. Rejuvenation Res 2009; 12(4):237-238.

de Grey ADNJ. Aging: can piecemeal preventative maintenance work? World Stem Cell Report 2009:101-103.

de Grey ADNJ. The Einsteinian imperative. Rejuvenation Res 2009; 12(6):383-385.

de Grey ADNJ. Hype and anti-hype in academic biogerontology research. Rejuvenation Res 2010; 13(2-3):137-138.

Rae MJ, Butler RN, Campisi J, de Grey ADNJ, Finch CE, Gough M, Martin GM, Vijg J, Perrott KM, Logan BJ. The demographic and biomedical case for late-life interventions in aging. Sci Transl Med 2010;2(40):40cm21.

de Grey ADNJ. Reaping the longevity dividend in time: biogerontology heavyweights advocate seeking late-onset interventions against aging. Rejuvenation Res 2010;13(4):383-385.

de Grey ADNJ. Biology: a field in need of a community culture. Rejuvenation Res 2010;13(5):507-508.

de Grey ADNJ. European Council of Art: a new initiative to combine both fields and cultures in the pursuit of visionary goals.Rejuvenation Res 2010;13(6):629-630.

de Grey ADNJ. Premature deaths close to home. Rejuvenation Res 2011;14(1):1-2.

de Grey ADNJ. Silver linings of oversimplification. Rejuvenation Res 2011;14(2):109-110.

de Grey ADNJ. The evolution of dogma. Rejuvenation Res 2011;14(3):239-240.

de Grey ADNJ. Progress... but speed is of the essence. Rejuvenation Res 2011;14(4):351-352.

de Grey ADNJ. When in doubt, maximize your options. Rejuvenation Res 2011;14(5):467-468.

de Grey ADNJ. The many-level value of proofs of concept. Rejuvenation Res 2012;15(2):121-122.

Zealley BJ, de Grey ADNJ. Cancers co-opt cohabitants' catabolism: Autophagy and senescence in the tumor stroma. Cell Cycle 2012;11(12):2230.

de Grey ADNJ. Biological versus medical limits on aging: a distinction we must not elide. Rejuvenation Res 2012;15(3):255-256.

de Grey ADNJ. What's really delaying the defeat of aging? Rejuvenation Res 2012;15(4):347-348.

de Grey ADNJ. The future dominance of combination therapies: implications for today's medical research. Rejuvenation Res 2012;15(5):443-444.

de Grey ADNJ. Dilemmas of discussing dramatic developments. Rejuvenation Res 2012;15(6):543-544.


- Books:

de Grey ADNJ.  The mitochondrial free radical theory of aging. Austin, TX: Landes Bioscience, 1999, 212pp, hardcover (ISBN 1-57059-564-X).

de Grey ADNJ, Rae M. Ending Aging: The rejuvenation biotechnologies that could reverse human aging in our lifetime. New York, NY: St. Martin’s Press, 2007, 416pp, hardcover (ISBN 0-312-36706-6). Paperback edition: St. Martin’s Press, 2008, 448pp (ISBN 0-312-36707-4).


- Book chapters:

de Grey ADNJ. Are those 13 proteins really unimportable? In: From Symbiosis to Eukaryotism - Endocytobiology VII (E. Wagner et al., eds.), Geneva University Press, 1999, pp. 489-502.

de Grey ADNJ. Mitochondrial mutations in vertebrate aging. In: Oxidative stress and aging: advances in basic science, diagnostics, and intervention (R.G. Cutler and H. Rodriguez, eds.), World Scientific Publishing, 2002, pp. 437-451.

de Grey ADNJ. Mechanisms underlying the age-related accumulation of mutant mitochondrial DNA. In: Genetics of mitochondrial diseases (I.J. Holt, ed.), Oxford University Press, 2003, pp. 247-275.

de Grey ADNJ. An engineer’s approach to the development of real anti-aging medicine. In: The Fountain of Youth: Ethical, Religious, and Existential Perspectives on a Biomedical Goal (S.G. Post and R.H. Binstock, eds.), Oxford University Press, 2004, pp. 249-267.

de Grey ADNJ. The war on aging. In: The Scientific Conquest of Death (B.J. Klein et al., eds.), Libros en Red, 2004, pp. 17-29.

de Grey ADNJ. Foreseeable and more distant rejuvenation therapies. In: Aging Interventions and Therapies (S.I.S. Rattan, ed.), World Scientific, 2005, pp. 379-395.

de Grey ADNJ. The foreseeability of real anti-aging medicine. In: Anti-Aging Medical Therapeutics, volume 7 (R. Klatz, ed.), American Academy for Anti-Aging Medicine, 2005, pp. 59-68.

de Grey ADNJ. Lysosomal enhancement with microbial hydrolases: a novel strategy for removing protein aggregates. In: New Trends in Alzheimer and Parkinson Disorders: ADPD 2005 (A. Fisher et al., eds.), Medimond, 2005, pp. 51-54.

de Grey ADNJ. Gene therapy. In: Encyclopedia of Aging, 4th Edition (R. Schulz et al., eds.), Springer, 2006, pp. 439-443.

de Grey ADNJ. Strategie per un invecchiamento transcurabile ingegnerizzato. In: Alterando il destino dell’umanità (P. Donghi, ed.), Proceedings of the 2005 Spoletoscienza conference, Laterza & Figli, 2006, pp. 49-63.

de Grey ADNJ. Defeat  of aging - utopia or foreseeable scientific reality? In: The Future of Life and the Future of our Civilization (V. Burdyuzha, ed.), Springer, 2006, pp. 277-290.

de Grey ADNJ. Rejuvenating neurons and glia with microbial enzymes. In: Interaction Between Neurons and Glia in Aging and Disease (J.O. Malva et al., eds.), Springer, 2007, pp. 503-510.

de Grey ADNJ. Endless Youth. In: Eindeloos, 22ste Almanak der Groningse Farmaceutische Studenten Vereniging “Pharmaciae Sacrum” (I. Dubbelboer et al., eds.), 2008, pp. 22-27.

de Grey ADNJ. Postponing aging: re-identifying the experts? In: Unnatural Selection: the Challenges of Engineering Tomorrow’s People (P. Healey and S. Rayner, eds.), 2008, pp. 89-94.

de Grey ADNJ. Radical life extension: technological aspects. In: Religion and the Implications Of Radical Life Extension (D.F. Maher and C. Mercer, eds.), Palgrave Macmillan, 2009, pp. 13-24.

de Grey ADNJ. WILT: necessity, feasibility, affordability. In: The Future of Aging: Pathways to Human Life Extension (G. Fahy et al., eds.), Springer, 2010, pp. 667-685.

de Grey ADNJ. Physical resilience and aging: correcting the Tithonus error and the crème brulée error. In: New Frontiers of Resilient Aging: Life-Strengths and Well-Being in Late Life (P.S. Fry and C.L.M. Keyes, eds.), 2010, pp. 90-103.

de Grey ADNJ. Prospects for the biomedical postponement of aging: technical context for a theological debate. In: The Routledge Companion to Religion and Science (J.W. Haag, G.R. Peterson and M.L. Spezio, eds.), 2011, pp. 531-539.

de Grey ADNJ. Living to 100 and maybe much longer: the engineering and biotechnology of life-extension medicine and when it may arrive. In: Proceedings of the 6th International Conference on Intelligent Processing and Manufacturing of Materials, (J.A. Meech, ed.), 2012, in press.

de Grey ADNJ. Is the quest to defeat aging ethical? In: Ethical Futures (S. Wint, ed.), 2012, in press.

de Grey ADNJ. Consciousness in the context of radical life extension. In: Holistic Consciousness: Cognition, Memory, Individuality, Free Will, Zombies, Life, Death, Immortality and the Immune System (N. Bauer, ed.), 2012, in press.

de Grey ADNJ. The future of regenerative medicine: the undoing of ageing. In: A Manual of Current Therapies in Regenerative Medicine (J.-T. Schantz, ed.), 2012, in press.


Invitations to speak:

- Seminars:

  University of Indiana, Bloomington, IN, USA, April 1997

  University of California, Riverside, CA, USA, February 1998

  University of Newcastle, UK, October 1998

  University of Linköping, Sweden, May 1999

  University of Marburg, Germany, July 1999

  ETH (Eidgenössische Technische Hochschule), Zurich, Switzerland, September 1999

  MRC Laboratory of Molecular Biology, Cambridge, UK, November 1999

  MRC Dunn Human Nutrition Unit, Cambridge, UK, January 2000

  University of Washington, Seattle, WA, USA, June 2000

  University of California, Irvine, CA, USA, October 2000

  University of California, Los Angeles, CA, USA, October 2000

  University of Southern California, Los Angeles, CA, USA, October 2000

  Kronos Longevity Research Institute, Phoenix, AZ, USA, February 2002

  University of Southampton, UK, April 2002

  Gray Cancer Institute, Northwood, UK, September 2002

  University of Newcastle, UK, February 2003

  University of California, Santa Barbara, CA, USA, March 2003

  Unilever Research and Development, Colworth, UK, April 2004

  University of Tampere, Finland, May 2004

  ExtroBritannia, Wolverton, UK, July 2004

  CCLRC Daresbury Laboratory, Warrington, UK, October 2004

  University of Birmingham, UK, November 2004

  University of Calgary, Calgary, AB, Canada, February 2005

  University of Alberta, Edmonton, AB, Canada, February 2005

  Baylor College of Medicine, Houston, TX, USA, March 2005

  Rice University, Houston, TX, USA, March 2005

  University of Texas Health Sciences Center, San Antonio, TX, USA, March 2005

  Café Scientifique, Edinburgh, UK, April 2005

  Hunter College, New York, NY, USA, April 2005

  Stanford University, Stanford, CA, USA, June 2005 (two talks)

  University of Utrecht, Utrecht, The Netherlands, September 2005

  London Regenerative Medicine Network, University College, London, UK, October 2005

  University of Cambridge Institute of Biotechnology, Cambridge, UK, December 2005

  Kronos Longevity Research Institute, Phoenix, AZ, USA, December 2005

  People Unlimited, Scottsdale, AZ, USA, December 2005

  Loma Linda University, Loma Linda, CA, USA, February 2006

  Faith, Reason and Community Forum, Loma Linda, CA, USA, February 2006

  Knutsford SciBAr, Knutsford, UK, February 2006

  Demos, London, UK, February 2006

  Sierra Sciences, Reno, NV, USA, February 2006

  Corven Corporate Finance, London, UK, March 2006

  University of Cambridge Medical Society, Cambridge, UK, March 2006

  University of Arizona, Tucson, AZ, USA, April 2006

  Arizona State University Biodesign Institute, Tempe, AZ, USA, April 2006

  Yale University, New Haven, CT, USA, May 2006

  Junto, New York, NY, USA, June 2006

  BUPA, London, UK, September 2006

  University of Michigan, Ann Arbor, MI, USA, October 2006

  ExtroBritannia, London, UK, October 2006

  The Pensions Net Work, Maidstone, UK, December 2006

  University of Heidelberg, Heidelberg, Germany, January 2007

  Cambridge University Student Pugwash Society, Cambridge, UK, February 2007

  Buck Institute for Age Research, Novato, CA, USA, March 2007

  University of Hohenheim, Stuttgart, Germany, April 2007

  Google, Mountain View, CA, USA, May 2007

  Royal Society of Arts, London, UK, June 2007

  Google, Mountain View, CA, USA, June 2007

  San Francisco Future Salon, Palo Alto, CA, USA, July 2007

  Cambridge Hi-tech Association of Small Enterprises, Cambridge, UK, August 2007

  British Humanist Association, London, UK, August 2007

  University of California Berkeley, Berkeley, CA, USA, October 2007

  Oxford University Science Society, Oxford, UK, October 2007

  Sangamo Sciences, Richmond, CA, USA, October 2007

  Milner Society (Queens’ College Cambridge), Cambridge, UK, November 2007

  Case Western Reserve University, Cleveland, OH, USA, November 2007 (two talks)

  Royal College of Arts, London, UK, November 2007

  Google, Mountain View, CA, USA, December 2007

  Bangor SciBAr, Bangor, UK, January 2008

  Dana Centre, London, UK, January 2008

  Trinity College Science Society, Cambridge, UK, January 2008

  University of Pennsylvania, Philadelphia, PA, February 2008

  New York University, New York, NY, USA, February 2008

  Cass Business School, London, UK, February 2008

  Cambridge Young Solicitors and Professionals Group, Cambridge, UK, February 2008

  St Paul’s School, London, UK, March 2008

  Spirit of the Senses, Phoenix, AZ, USA, March 2008

  University of Wisconsin, Madison, WI, USA, April 2008

  Orlando Science Center, Orlando, FL, USA, May 2008

  Proctor & Gamble workshop “Future thinking”, New York, NY, USA, June 2008

  University of Alberta, Edmonton, Canada, June 2008

  The Pensions Net Work, Hurley, UK, June 2008

  Toronto Public Library Network, Toronto, Canada, June 2008

  ExtroBritannia, London, UK, July 2008

  University of Hull, Hull, UK, July 2008

  Café Scientifique, Lichfield, UK, July 2008

  The Motley Fool, Alexandria, MD, USA, July 2008

  Google, New York, NY, USA, July 2008

  UCLA student group “POLIS”, Los Angeles, CA, USA, August 2008

  NuSkin, Provo, UT, USA, August 2008

  NASA Ames Research Center, Mountain View, CA, UA, August 2008

  Think Tank der Bank Sarasin & Cie, Zurich, Switzerland, September 2008

  Kronos Longevity Research Institute, Scottsdale, AZ, USA, September 2008

  University of Ghent, Ghent, Belgium, October 2008

  Jewish Community Center of San Francisco, San Francisco, CA, USA, October 2008

  Stanford Transhumanist Association, Palo Alto, CA, USA, October 2008

  The Nature Debate, London, UK, November 2008

  Spirit of the Senses, Phoenix, AZ, USA, December 2008

  HEALES, Narbonne, France, January 2009

  Caixa Foundation, Lleida, Spain, January 2009

  Caixa Foundation, Barcelona, Spain, January 2009

  Caixa Foundation, Madrid, Spain, January 2009

  University of Palma, Palma, Mallorca, January 2009

  Caixa Foundation, Tarragona, Spain, January 2009

  Kent State University, Kent, OH, USA, February 2009

  University of Manchester, Manchester, UK, February 2009

  University of Liverpool, Liverpool, UK, February 2009

  Moscow State University, Moscow, Russia, March 2009

  University of Westminster, London, UK, March 2009

  University of Ghent, Ghent, Belgium, March 2009

  Bank Sarasin & Cie, Interlaken, Switzerland, April 2009

  Queen Mary University of London, London, UK, April 2009

  Smart Life Forum, Palo Alto, CA, USA, May 2009

  Commonwealth Club, San Francisco, CA, USA, May 2009

  University of Leicester, Leicester, UK, June 2009

  Singularity University, Mountain View, CA, USA, July 2009

  Summer Science Program, Ojai, CA, USA, July 2009

  Skeptics in the Pub, London, UK, August 2009

  Kings Fund, London, UK, September 2009

  New York Academy of Sciences, New York, NY, USA, September 2009

  Silicon Valley Transhumanist Meetup, Palo Alto, CA, USA, September 2009

  The Commonwealth Medical College, Scranton, PA, USA, September 2009

  UK Transhumanist Association, London, UK, October 2009

  Oxford University Science Society, Oxford, UK, October 2009

  Karolinska Institute, Stockholm, Sweden, October 2009 (two talks)

  Osher Lifelong Learning Institute, Saginaw, MI, USA, November 2009

  NHS Workforce Review Team, Winchester, UK, December 2009

  Haberdashers’ Aske’s Boys’ School, Elstree, UK, January 2010

  Wellcome Trust, London, UK, January 2010

  Staffordshire University, Stafford, UK, March 2010

  Omega Institute, Rhinebeck, NY, USA, October 2010


- National and international meetings:


  Endocytobiology VII, Freiburg, Germany, April 1998

  4th European Congress of Gerontology, Berlin, Germany, July 1999

  Euromit 4, Cambridge, UK, September 1999

  Symposium: stress and ageing in animals and microorganisms, Porto, Portugal, June 2000

  BBSRC Science of Ageing symposium, Manchester, UK, January 2001

  O2SA (Oxidative Stress in Aging), 2nd Conference, Maui, HI, USA, April 2001

  International Association of Biomedical Gerontology, 9th Congress, Vancouver, Canada, June 2001

  27th Meeting of FEBS, Lisbon, Portugal, July 2001

  Warburg Society, inaugural conference, Point Clear, LA, USA, October 2001

  6th International Conference on Plasma Membrane Redox Systems, Ravenna, Italy, March 2002

  Mechanisms of Caloric Restriction conference, Bandera, TX, USA, October 2002

  3rd European Congress of Biogerontology pre-meeting, San Miniato, Italy, November 2002

  5th Alcor Conference on Extreme Life Extension, Newport Beach, CA, USA, November 2002

  Foresight Institute 2003 Senior Associates Gathering, Palo Alto, CA, USA, May 2003

  Transvision 2003, Yale University, New Haven, CT, USA, June 2003

  CMIB symposium “Projecting Future Mortality”, Edinburgh, UK, October 2003

  Pop!Tech 2003 conference “Sea Change”, Camden, ME, USA, October 2003

  7th International Conference on Plasma Membrane Redox Systems, Asilomar, CA, USA, April 2004

  Pittsburgh symposium “Frontiers in Human Embryonic Stem Cells”, Pittsburgh, PA, USA, May 2004

  3rd Medical and Aesthetic Procedures Meeting, Barcelona, Spain, June 2004

  3rd Asia Pacific Conference and Exhibition on Anti-Ageing Medicine, Singapore, June 2004

  Transvision 2004, Toronto, Canada, August 2004

  1st Anti-Ageing Conference London, London, UK, September 2004

  German Society of Gerontology and Geriatrics, 7th Congress, Hamburg, Germany, October 2004

  5th Neurobiology of Aging symposium, San Diego, CA, USA, October 2004

  Integrative Medical Therapeutics for Anti-Aging conference, Las Vegas, NV, USA, October 2004

  EMBL/EMBO conference on Science and Society, Heidelberg, Germany, November 2004

  Futurshow 3004, Milan, Italy, November 2004

  12th Annual World Congress on Anti-Aging Medicine, Las Vegas, NV, USA, December 2004

  BIOMEDEX 2005, Montreal, Canada, March 2005

  IHEU-Appignani Humanist Center for Bioethics, 1st Conference, New York, NY, USA, April 2005

  2nd “The future of life and the future of our civilization” symposium, Frankfurt, Germany, May 2005

  2nd Anti-Ageing Conference London, London, UK, May 2005

  Workshop on Models in the Biodemography of Aging, Berkeley, CA, USA, June 2005

  FENS Summer School, Coimbra, Portugal, July 2005

  TEDGLOBAL (Technology, Entertainment, Design) conference, Oxford, UK, July 2005

  17th SpoletoScienza conference “Altering Human Destiny”, Spoleto, Italy, July 2005

  Eris Society, 25th annual conference, Aspen, CO, USA, August 2005

  International Association of Biomedical Gerontology, 11th Congress, Aarhus, Denmark, August 2005

  10th Future of Health Technology Summit, Cambridge, MA, USA, September 2005

  4th RNAO International Elder Care Conference, Toronto, Canada, September 2005

  Conservative Party Conference 2005, Blackpool, UK, October 2005

  Louis Vuitton Moet Hennessy symposium on mitochondria, Paris, France, October 2005

  European Society for Cosmetic & Aesthetic Dermatology Society Mtg., London, UK, October 2005

  Foresight Institute 2005 Vision Weekend, San Francisco, CA, USA, October 2005

  Immortality Institute, 1st Conference, Atlanta, GA, USA, November 2005

  58th Annual Meeting, Gerontological Society of America, Orlando, FL, USA, November 2005

  Nordic Exceptional Trendshop 2005, Copenhagen, Denmark, November 2005

  22nd Human Genetics Commission Plenary Meeting, November 2005

  9th Goodenough conference ‘Age & Ageing: Just the Concern of the Old?’, London, UK, Dec. 2005

  AAAS Annual Meeting, St. Louis, MO, USA, February 2006

  TED (Technology, Entertainment, Design) 2006, Monterey, CA, USA, February 2006

  Vodafone Foundation symposium, Madrid, Spain, March 2006

  1st World Forum on Science and Civilisation, Oxford, UK, March 2006

  1st Berkeley Stem Cell Retreat, Asilomar, CA, USA, March 2006

  Calorie Restriction Society, 4th Conference, Tucson, AZ, USA, April 2006

  Human Enhancement Technologies and Human Rights, Stanford, CA, USA, May 2006

  Transvision 2006, Helsinki, Finland, August 2006                                                                                   

  1st ENHANCE Workshop on the Extension of Lifespan, Cesano Maderno, Italy, September 2006

  5th European Congress of Biogerontology, Istanbul, Turkey, September 2006

  2nd Women’s Forum for the Economy and Society, Deauville, France, October 2006

  6th Alcor Conference on Extreme Life Extension, Scottsdale, AZ, USA, October 2006

  Living Well to 100, 2nd Conference, Boston, USA, November 2006

  3rd World Mystery Forum, Interlaken, Switzerland, November 2006

  American Academy of Religion, 2006 Annual Meeting, Washington, DC, USA, November 2006

  Rome Science Festival, Rome, Italy, January 2007

  London Technology Network: Stem Cells and Regenerative Medicine, London, UK, March 2007

  Edmonton Aging Symposium, Edmonton, Canada, March 2007

  UBS Global Investors’ Forum, London, UK, April 2007

  German Society of Anti-Aging Medicine, 7th Conference, Munich, Germany, May 2007

  2nd ENHANCE Workshop on the Extension of Lifespan, Cesano Maderno, Italy, May 2007

  International Association of Biomedical Gerontology, 12th Congress, Spetses, Greece, May 2007

  6th Cheltenham Science Festival, Cheltenham, UK, June 2007

  6th Intl. Conf. on Intelligent Processing and Manufacturing of Materials, Salerno, Italy, June 2007

  Oxford Conference on Genetics and Justice, Oxford, UK, July 2007

  National Council for Palliative Care, 15th Anniversary Symposium, London, UK, July 2007

  CEPS symposium “Europe facing ageing - economic perspectives”, Prague, Czech Rep., July 2007

  Institute for Ethics and Emerging Technologies symposium, Chicago, IL, USA, July 2007

  Transvision 2007, Chicago, IL, USA, July 2007

  Secret Garden Party, Kings Ripton, UK, July 2007

  4th Anti-Ageing Conference London, London, UK, September 2007

  12th Future of Health and Technology Institute Conference, Cambridge, MA, USA, September 2007

  Gadgetoff, New York, NY, USA, September 2007

  7th Alcor Conference on Extreme Life Extension, Scottsdale, AZ, USA, October 2007

  3rd European Anti-Aging Congress, Paris, France, October 2007

  8th Asia/Oceania Regional Congress of Gerontology & Geriatrics, Beijing, China, October 2007

  American Academy of Religion, 2007 Annual Meeting, San Diego, CA, USA, November 2007

  Hewitt’s Local Government Pension Service symposium, London, UK, January 2008

  Future of GI Health Summit, Boston, MA, USA, January 2008

  BIL 2008, Monterey, CA, USA, March 2008

  Center for Bioethics & Human Dignity, 1st Annual Phoenix Conf., Phoenix, AZ, USA, March 2008

  Anti-Aging Medicine World Congress, Paris, France, April 2008

  34th Asilomar Microcomputer Workshop, Asilomar, CA, USA, April 2008

  Seoul Digital Forum, Seoul, Korea, May 2008

  ERBI’s 10th Annual BioPartnering Exchange, Hinxton, UK, May 2008

  Hewlett-Packard Worldwide Health and Life Sciences Symposium, Orlando, FL, USA, May 2008

  Italian Academy of Anti-Aging Medicine, 3rd National Congress, May 2008

  Life Link conference, Gleneagles, UK, May 2008

  Thinking Digital, Gateshead, UK, May 2008

  Anti-Ageing Skin Care Conference 2008, London, UK, June 2008

  IdeaCity08, Toronto, Canada, June 2008

  BioBarCamp, Mountain View, CA, USA, August 2008

  SciFoo, Mountain View, CA, USA, August 2008

  European Congress on Anti-Aging and Aesthetic Medicine, Dusseldorf, Germany, September 2008

  13th Future of Health and Technology Institute Conference, Cambridge, MA, USA, September 2008

  International Workshop on postponing ageing, Tallinn, Estonia, October 2008

  2008 Conference of the Libertarian Alliance, London, UK, October 2008

  4th European Futurists Conference, Lucerne, Switzerland, October 2008

  The Future, Hamburg, Germany, October 2008

  Summit on the Global Agenda, Dubai, November 2008

  Changing the World, Toronto, Canada, November 2008

  Convergence ‘08, Mountain View, CA, USA, November 2008

  ASU workshop on the systems biology of aging, Tempe, AZ, USA, December 2008

  BIL 2009, Monterey, CA, USA, February 2009

  Workshop on Anti-Aging Medicine, Volterra, Italy, March 2009

  4th World Congress of Regenerative Medicine, Bangkok, Thailand, March 2009

  Eisai Symposium in Neuroscience, London, UK, April 2009

  International Association of Biomedical Gerontology, 13th Congress, Montreal, Canada, May 2009

  Metanexus Conference, Phoenix, AZ, USA, July 2009

  Life Aid, Brussels, Belgium, September 2009

  World Stem Cell Summit, Baltimore, MD, USA, September 2009

  Glenn Foundation Symposium on the Biology of Aging, Cambridge, MA, USA, September 2009

  14th Future of Health Technology Summit, Cambridge, MA, USA, September 2009

  IQPC Life Settlements Summit, New York, NY, USA, October 2009

  Singularity Summit, New York, NY, USA, October 2009

  2nd European Conference on Anti-Aging and Aesthetic Medicine, Mainz, Germany, October 2009

  Women’s Forum Global Meeting, Deauville, France, October 2009

  Brocher Foundation symposium, Geneva, Switzerland, October 2009

  TED MED 2009, San Diego, CA, USA, October 2009

  BIL:PIL, San Diego, CA, USA, October 2009

  Manhattan Beach Project, Manhattan Beach, CA, USA, November 2009

  UK Society of Cosmetic Scientists, London, UK, December 2009

  Systems biology of aging, 2nd annual conference, Baltimore, MD, USA, December 2009

  LIFT, Geneva, Switzerland, May 2010

  Anti-Aging Medicine conference, Lisbon, Portugal, May 2010

  American Aging Association, Portland, OR, USA, June 2010

  World Future Society, Boston, MA, USA, July 2010

  European Respiratory Society, Barcelona, Spain, September 2010

  Our Common Future, Essen, Germany, November 2010

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